subgenus

Doratulina (Doratulina) Melichar, 1903

Habitus, dorsal view: !! display not done !! on Doratulina (Doratulina) Melichar, 1903: Doratulina dmitrievi, male, dorsal habitus (INHS). (Observation).

Habitus, lateral view: !! display not done !! on Doratulina (Doratulina) Melichar, 1903: Doratulina dmitrievi, female, lateral habitus (INHS). (Observation).

Pygofer, lateral view: !! display not done !! on Doratulina (Doratulina) Melichar, 1903: Doratulina dmitrievi, pygofer, laterally (INHS). (Observation).

Subgenital plate: !! display not done !! on Doratulina (Doratulina) Melichar, 1903: Doratulina dmitrievi, subgenital plates and valve, ventrally (INHS). (Observation).

Aedeagus, ventral view: !! display not done !! on Doratulina (Doratulina) Melichar, 1903: Doratulina dmitrievi, aedeagus, caudally (INHS). (Observation).

Aedeagus, lateral view: !! display not done !! on Doratulina (Doratulina) Melichar, 1903: Doratulina dmitrievi, aedeagus, ventrally (INHS). (Observation).

Connective: !! display not done !! on Doratulina (Doratulina) Melichar, 1903: Doratulina dmitrievi, connective and stylus, ventrally (INHS). (Observation).

Pygofer, dorsal view: !! display not done !! on Doratulina (Doratulina) Melichar, 1903: Doratulina dmitrievi, pygofer, dorsally (INHS). (Observation).

Ovipositor: !! display not done !! on Doratulina (Doratulina) Melichar, 1903: Doratulina dmitrievi, female sternite VII, ventrally (INHS). (Observation).

Descendants and synonyms

Nomenclature (57)

Doratulina Melichar, 1903b: 198. (valid in Emeljanov, 1999e: 605)
type species by original designation Doratulina jocosa Melichar, 1903
Anemochrea Kirkaldy, 1906c: 329.
type species by original monotypy Anemochrea mitis Kirkaldy, 1906
... Show all ... (53)

Phrynophyes Kirkaldy, 1906c: 327.
type species by original designation Phrynophyes phrynophyes Kirkaldy, 1906
Phrynophryes [sic] Kirkaldy, 1906 in Perkins, 1906b: 495.
Phrynophryes [sic] Kirkaldy, 1906 (misspelling of Phrynophyes Kirkaldy, 1906) in Perkins, 1906b: 495.
Doratula [sic] Melichar, 1903 in Melichar, 1908b: 14.
Arya Distant, 1908g: 338.
type species by original designation Arya rubrolineata Distant, 1908
Doratula [sic] Melichar, 1903 (misspelling of Doratulina Melichar, 1903) in Melichar, 1908b: 14.
Bituitus Distant, 1918b: 70.
type species by original designation Bituitus projectus Distant, 1918
Campbellinella Distant, 1918b: 69.
type species by original designation Campbellinella illustrata Distant, 1918
Nandidrug Distant, 1918b: 74.
type species by original designation Nandidrug speciosum Distant, 1918
Paivanana Distant, 1918b: 95.
type species by original designation Typhlocyba indra Distant, 1908
Paternus Distant, 1918b: 71.
type species by original designation Paternus pusanus Distant, 1918
Volusenus Distant, 1918b: 72.
type species by original designation Volusenus lahorensis Distant, 1918
Bella Singh-Pruthi, 1930a: 44.
type species by original designation Bella apicalis Singh-Pruthi, 1930
Sunda Singh-Pruthi, 1936a: 112.
type species by original designation Sunda ribeiroi Singh-Pruthi, 1936
Pseudaconura Linnavuori, 1952b: 182.
type species by original designation Pseudaconura luxorensis Linnavuori, 1952
Coratulina [sic] Melichar, 1903 in Esaki & Ito, 1954a: 110.
Malagasiella Evans, 1954a: 110.
type species by original designation Malagasiella minima Evans, 1954
Coratulina [sic] Melichar, 1903 (misspelling of Doratulina Melichar, 1903) in Esaki & Ito, 1954a: 110.
Umesaona Ishihara, 1961a: 246.
type species by original designation Umesaona asiatica Ishihara, 1961
Doratulina Melichar, 1903 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Anemochrea Kirkaldy, 1906 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Arya Distant, 1908 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Bella Singh-Pruthi, 1930 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Bituitus Distant, 1918 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Campbellinella Distant, 1918 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Nandidrug Distant, 1918 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Paivanana Distant, 1918 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Paternus Distant, 1918 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Phrynophyes Kirkaldy, 1906 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Pseudaconura Linnavuori, 1952 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Sunda Singh-Pruthi, 1936 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Umesaona Ishihara, 1961 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Volusenus Distant, 1918 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Emeljanov, 1962b: 393.
Dortulina [sic] Melichar, 1903 in Anonymous, 1964: 282.
Dortulina [sic] Melichar, 1903 (misspelling of Doratulina Melichar, 1903) in Anonymous, 1964: 282.
Umesdena [sic] Ishihara, 1961 in Vilbaste, 1965a: 10.
Anemochrea Kirkaldy, 1906 (subjective synonym of Doratulina Melichar, 1903) in Vilbaste, 1965a: 10.
Arya Distant, 1908 (subjective synonym of Doratulina Melichar, 1903) in Vilbaste, 1965a: 10.
Bella Singh-Pruthi, 1930 (subjective synonym of Doratulina Melichar, 1903) in Vilbaste, 1965a: 10.
Bituitus Distant, 1918 (subjective synonym of Doratulina Melichar, 1903) in Vilbaste, 1965a: 10.
Campbellinella Distant, 1918 (subjective synonym of Doratulina Melichar, 1903) in Vilbaste, 1965a: 10.
Nandidrug Distant, 1918 (subjective synonym of Doratulina Melichar, 1903) in Vilbaste, 1965a: 10.
Paivanana Distant, 1918 (subjective synonym of Doratulina Melichar, 1903) in Vilbaste, 1965a: 10.
Paternus Distant, 1918 (subjective synonym of Doratulina Melichar, 1903) in Vilbaste, 1965a: 10.
Phrynophyes Kirkaldy, 1906 (subjective synonym of Doratulina Melichar, 1903) in Vilbaste, 1965a: 10.
Sunda Singh-Pruthi, 1936 (subjective synonym of Doratulina Melichar, 1903) in Vilbaste, 1965a: 10.
Umesaona Ishihara, 1961 (subjective synonym of Doratulina Melichar, 1903) in Vilbaste, 1965a: 10.
Volusenus Distant, 1918 (subjective synonym of Doratulina Melichar, 1903) in Vilbaste, 1965a: 10.
Umesdena [sic] Ishihara, 1961 (misspelling of Umesaona Ishihara, 1961) in Vilbaste, 1965a: 10.
Pseudaconura Linnavuori, 1952 (subjective synonym of Doratulina Melichar, 1903) in Nast, 1972a: 354.
Doratulina Melichar, 1903 (classified as Stirellini Emeljanov, 1966) in Hamilton, 1975b: 481.
Malagasiella Evans, 1954 (subjective synonym of Doratulina Melichar, 1903) in Zahniser & Nielson, 2012a: 49.
Doratulina Melichar, 1903 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Duan, Zhang & Dietrich, 2019a: 205.

Malagasiella Evans, 1954 (subjective synonym of Athysanus (Stirellus) Osborn & Ball, 1902) in Duan, Zhang & Dietrich, 2019a: 205.
Malagasiella Evans, 1954 (subjective synonym of Doratulina Melichar, 1903) in Dmitriev, 2024

Type

Doratulina jocosa Melichar, 1903 type species by original designation of Doratulina (Doratulina) Melichar, 1903

Gender, form, and etymology

Etymology:

Doratura + -ina (related to, wife of)

Nomenclature references (22)

Anonymous (1964) Arocephalus monticulinus. The Zoological Record, 101(13), 282.
Distant, W.L. (1908g) Rhynchota. IV. Homoptera and appendix (Pt.). In The fauna of British India, including Ceylon and Burma. London: Taylor & Francis. pp. 1–419. [XV + 501 pp.]
... Show all ... (18)

Distant, W.L. (1918b) Rhynchota. VII. Homoptera: appendix. Heteroptera: addenda. In A.E. Shipley & G.A.K. Marshall (Eds.). The fauna of British India, including Ceylon and Burma. Taylor & Francis, London. Vol. 7, pp. i-viii + 1–210.

Dmitriev, D.A. (2024) Personal communication.

Duan, Y.-N., Zhang, Y.-T. & Dietrich, C.H. (2019a) Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records. Zootaxa, 4603(2), 201–240. https://doi.org/10.11646/zootaxa.4603.2.1

Emeljanov, A.F. (1962b) New tribes of leafhoppers of the subfamily Euscelinae (Auchenorrhyncha. Cicadellidae). [Новые трибы цикадок из подсем. Euscelinae (Auchenorrhyncha, Cicadellidae)]. Entomologicheskoe Obozrenie, 41(2), 388–397. [in Russian; translated into English in Entomological Review, Washington, 41(2), 236–240]

Emeljanov, A.F. (1999e) A key to genera of the subfamily Deltocephalinae s. l. (Homoptera, Cicadellidae) from Kazakhstan, Middle Asia and Mongolia with description of new genera and subgenera. [Определительная таблица родов подсем. Deltocephalinae s. l. (Homoptera, Cicadellidae) Казахстана, Средней Азии и Монголии с описанием новых родов и подродов]. Entomologicheskoe Obozrenie, 78(3), 589–609. [in Russian; translated into English in Entomological Review, Washington, 79(5), 547–562]

Esaki, T. & Ito, S. (1954a) A tentative catalogue of Jassoidea of Japan, and her adjacent territories. Japan Society for Promotion of Science. Ueno Park, Tokyo, 315 pp.

Evans, J.W. (1954a) Les Cicadellidae de Madagascar (Homopteres). Mémoires de l'Institut Scientifique de Madagascar, Série E Entomologie, 4, 87–137.

Hamilton, K.G.A. (1975b) Review of the tribal classification of the leafhopper subfamily Aphrodinae (Deltocephalinae of authors) of the Holarctic region (Rhynchota: Homoptera: Cicadellidae). The Canadian Entomologist, 107, 477–498.

Ishihara, T. (1961a) Homoptera of Southeast Asia collected by the Osaka City University Biological Expedition to Southeast Asia 1957-1958. In T. Kira & T. Umesao (Eds.). Nature & Life in Southeast Asia. Fauna and Flora Research Society, Kyoto, Japan. Vol. 1, pp. 225–257.

Kirkaldy, G.W. (1906c) Leaf-hoppers and their natural enemies. (Pt. IX. Leaf-hoppers-Hemiptera). Report of work of the Experiment Station of the Hawaiian Sugar Planters' Association. Entomological series, 1(9), 271–479.

Linnavuori, R.E. (1952b) Studies on some Palearctic [sic!] Hemiptera. Annales Entomologici Fennici, 18(4), 181–187.

Melichar, L. (1903b) Homopteren-Fauna von Ceylon. In Verlag von Felix L. Dames. Berlin. pp. i-iv, 1-248. Verlag von Felix L. Dames. Berlin. iv + 248 pp. + 6 pls.

Melichar, L. (1908b) Nové rody a druhy Homopter z východni Afriky. Časopis České Společnosti Entomologické = Acta Societatis Entomologicae Bohemiae, 5, 1–15.

Nast, J. (1972a) Palaearctic Auchenorrhyncha (Homoptera). An annotated check list. Polish Scientific Publishers, Warszawa, 550 pp.

Osborn, H. & Ball, E.D. (1902a) A review of the North American species of Athysanus (Jassidae). The Ohio Naturalist, 2, 231–256.

Perkins, R.C.L. (1906b) Leaf-hoppers and their natural enemies (Pt. X. Dryinidae, Pipunculidae). Supplementary. Bulletin. Hawaiian Sugar Planters' Association Experiment Station. Division of Entomology., 1(10), 483–499.

Singh-Pruthi, H. (1930a) Studies on Indian Jassidae (Homoptera). Part I. Introductory and description of some new genera and species. Memoirs of the Indian Museum, 11, 1–68 + pls. I-V.

Singh-Pruthi, H. (1936a) Studies on Indian Jassidae (Homoptera). Part III. Descriptions of some new genera and species, with first records of some known species from India. Memoirs of the Indian Museum, 11, 101–131 + pls. VIII + IX.

Vilbaste, J. (1965a) On the Genus Aconura Leth. (Homoptera, Jassidae). Notulae Entomologicae, 45, 3–12.
Zahniser, J.N. & Nielson, M.W. (2012a) An extraordinary new genus and three new species of Acostemmini (Hemiptera: Cicadellidae: Deltocephalinae) from Madagascar with comments on the morphology and classification of the tribe. Zootaxa, 3209, 28–52.

Stats

	Valid names
Rank	Extant	Fossil	Invalid	Total
subgenus	1	0	19	20
species	33	0	49	82