species

Amrasca (Sundapteryx) biguttula (Ishida, 1913)

indian cotton jassid; okra leafhopper; cotton leafhopper; two-spot cotton leafhopper

Habitus, dorsal view: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Habitus. (Observation).

Habitus, lateral view: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Lateral. (Observation).

Head: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Head. (Observation).

Head: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Head, female. (Observation).

Head: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Head, male. (Observation).

Head: !! display not done !! on Amrasca (Sundapteryx) punctata (Melichar, 1914): Head, pro-, and mesothorax. (Observation).

Subgenital plate: !! display not done !! on Amrasca (Sundapteryx) punctata (Melichar, 1914): genital plate. (Observation).

Head: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Head and thorax. (Observation).

Subgenital plate: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Subgenital plate. (Observation).

Aedeagus, ventral view: !! display not done !! on Amrasca (Sundapteryx) punctata (Melichar, 1914): Aedeagus, ventral view. (Observation).

Aedeagus, ventral view: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Aedeagus, ventral view. (Observation).

Stylus: !! display not done !! on Amrasca (Sundapteryx) punctata (Melichar, 1914): Stylus or paramere. (Observation).

Stylus: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Stylus. (Observation).

Forewing: !! display not done !! on Amrasca (Sundapteryx) punctata (Melichar, 1914): Forewing. (Observation).

Aedeagus, lateral view: !! display not done !! on Amrasca (Sundapteryx) punctata (Melichar, 1914): Aedeagus, lateral view. (Observation).

Forewing: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Forewing. (Observation).

Aedeagus, lateral view: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Aedeagus, lateral view. (Observation).

Connective: !! display not done !! on Amrasca (Sundapteryx) punctata (Melichar, 1914): Connective. (Observation).

Abdomen, sternal apodemes, ventral view: !! display not done !! on Amrasca (Sundapteryx) punctata (Melichar, 1914): Sternal apodemes. (Observation).

Abdomen, sternal apodemes, ventral view: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Abdomen, sternal apodemes, ventral view. (Observation).

Abdomen, sternal apodemes, ventral view: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Abdomen, sternal apodemes, ventral view (E. devastans, Distant, 1918). (Observation).

Pygofer appendages: !! display not done !! on Amrasca (Sundapteryx) punctata (Melichar, 1914): Pygofer ventral appendage. (Observation).

Pygofer appendages: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Pygofer appendages. (Observation).

Anal tube processes: !! display not done !! on Amrasca (Sundapteryx) punctata (Melichar, 1914): Anal tube. (Observation).

Anal tube processes: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Anal tube process. (Observation).

Other: !! display not done !! on Amrasca (Sundapteryx) biguttula (Ishida, 1913): Labels. (Observation).

Descendants and synonyms

Nomenclature (40)

Empoasca devastans Maxwell-Lefroy, 1906a: 108. (nomen nudum: no description)
Chlorita biguttula Ishida, 1913a: 1.
Lectotype; male; 129103e9-cf25-4aa7-9b46-23432dd65fa2; deposited at: Entomological Institute, Hokkaido University (EIHU); Japan: Okinawa
... Show all ... (36)

Chlorita biguttula Shiraki, 1913b: 96.
Lectotype; male; 129103e9-cf25-4aa7-9b46-23432dd65fa2; deposited at: Entomological Institute, Hokkaido University (EIHU); Japan: Okinawa
Zygina biguttula punctata Melichar, 1914b: 146.
Lectotype; adult; e680b040-48d7-48a1-898d-67522694a494; deposited at: Senckenberg Museum für Tierkunde, Dresden, Germany (SMTD); Indonesia: East Sumbawa, former Ost-Soembawa, Dompoe, 24-25.V.1927, Rensch
Empoasca bipunctata Schumacher, 1915b: 127.
Chlorita bimaculata Matsumura, 1916b: 393.
Empoasca devastans Distant, 1918b: 93.
Chlorita bimaculata Matsumura, 1916 (subjective synonym of Chlorita biguttula Ishida, 1913) in Matsumura, 1934b: 5.
Empoasca nigropunctata Merino, 1936a: 389.
Holotype; adult; a186afb3-9ae5-42e1-8cb5-0047f14c8cf5; deposited at: National Museum of the Philippines, Manila (MPMP); Philippines: Luzon, Rizal Province, Alabang, J. Valdez
Empoasca biguttula (Ishida, 1913) in Anonymous, 1936d: 1.
Empoasca devastens [sic] (Distant, 1918) in Cherian & Kylasam, 1938a: 76.
Empoasca devastens [sic] (Distant, 1918) (misspelling of Empoasca devastans Distant, 1918) in Cherian & Kylasam, 1938a: 76.
Chlorita bigutulla [sic] (Ishida, 1913) in Husain & Lal, 1940a: 124.
Chlorita bigutulla [sic] (Ishida, 1913) (misspelling of Chlorita biguttula Ishida, 1913) in Husain & Lal, 1940a: 124.
Empoasca uniguttata Jacobi, 1941a: 311.
Empoasca quadrinotatissima Dlabola, 1957b: 296.
Chlorita quadrinotatissima (Dlabola, 1957) in Dlabola, 1958c: 52.
Empoasca schumacheri Metcalf, 1968a: 353.
Empoasca devastans Maxwell-Lefroy, 1906 (unavailable or invalid, linked to Chlorita biguttula Ishida, 1913) in Metcalf, 1968a: 314.
Typhlocyba uniguttata (Jacobi, 1941) in Metcalf, 1968a: 905.
Chlorita bimaculata Matsumura, 1916 (subjective synonym of Chlorita biguttula Ishida, 1913) in Dworakowska, 1970h: 712.
Empoasca bipunctata Schumacher, 1915 (subjective synonym of Chlorita biguttula Ishida, 1913) in Dworakowska, 1970h: 712.
Empoasca devastans Distant, 1918 (subjective synonym of Chlorita biguttula Ishida, 1913) in Dworakowska, 1970h: 712.
Empoasca quadrinotatissima Dlabola, 1957 (subjective synonym of Chlorita biguttula Ishida, 1913) in Dworakowska, 1970h: 712.
Empoasca uniguttata Jacobi, 1941 (subjective synonym of Zygina biguttula punctata Melichar, 1914) in Dworakowska, 1970h: 712.
Sundapteryx biguttula biguttula (Ishida, 1913) in Dworakowska, 1970h: 712.
Sundapteryx biguttulaaaaa punctata (Melichar, 1914) in Dworakowska, 1970h: 712.
Amrasca devastans (Distant, 1918) in Ahmed & Samad, 1972b: 288.
Empoasca schumacheri Metcalf, 1968 (unnecessary replacement for Empoasca bipunctata Schumacher, 1915) in Linnavuori, 1975c: 616.
Amrasca biguttula biguttula (Ishida, 1913) in Dworakowska & Viraktamath, 1975a: 530.
Amrasca biguttulaaaaa punctata (Melichar, 1914) in Dworakowska, 1976b: 4.
Amrasca biguttula (Ishida, 1913) in Dworakowska, 1977i: 14.
Chlorita biguttula Shiraki, 1913 (synonymic homonym of Chlorita biguttula Ishida, 1913) in Sohi, 1983a: 53.
Empoasca nigropunctata Merino, 1936 (subjective synonym of Chlorita biguttula Ishida, 1913) in Sohi, 1983a: 52.
Empoasca biguttala [sic] (Ishida, 1913) in Li, Xu & Zhang, 2005: 147.
Empoasca biguttala [sic] (Ishida, 1913) (misspelling of Chlorita biguttula Ishida, 1913) in Li, Xu & Zhang, 2005: 147.
Amrasca (Amrasca) biguttula (Ishida, 1913) in Khatri, Rustamani, Ahmed, Sultana & Zaidi, 2013: 1264.
Empoasca uniguttata Jacobi, 1941 (subjective synonym of Chlorita biguttula Ishida, 1913) in Xu, Wang, Dietrich, Fletcher & Qin, 2017a: 363.

Zygina biguttula punctata Melichar, 1914 (subjective synonym of Chlorita biguttula Ishida, 1913) in Xu, Wang, Dietrich, Fletcher & Qin, 2017a: 363.
Amrasca (Sundapteryx) biguttula (Ishida, 1913) in Xu, Wang, Dietrich, Fletcher & Qin, 2017a: 362.

Gender, form, and etymology

Etymology:

Latin: two small spots

Nomenclature references (26)

Ahmed, M. & Samad, K. (1972b) Some new additions to the Typhlocybine fauna of East Pakistan. Pakistan Journal of Scientific and Industrial Research, 15(4-5), 285–290.
Anonymous (1936d) [Results of cotton cultivation experiments in Formosa.]. Bulletin. Department of Agriculture, Government Research Institute. Taihoku, Formosa, 122, 1–113.
... Show all ... (22)

Cherian, M.C. & Kylasam, M.S. (1938a) Jassids on cotton. Jour. Madras Agr., 26, 76–77.

Distant, W.L. (1918b) Rhynchota. VII. Homoptera: appendix. Heteroptera: addenda. In A.E. Shipley & G.A.K. Marshall (Eds.). The fauna of British India, including Ceylon and Burma. Taylor & Francis, London. Vol. 7, pp. i-viii + 1–210.

Dlabola, J. (1957b) Die Zikaden Afghanistans (Homopt. - Auchenorrhyncha) nach den Ergebnissen der von Herrn J. Klapperich in den Jahren 1952-1953 nach Afghanistan unternommennen Expedition. Mitteilungen der Münchner Entomologischen Gesellschaft, 47(5), 265–303.

Dlabola, J. (1958c) A reclassification of palaearctic Typhlocybinae (Homopt., Auchenorrh.). Acta Societatis Entomologicae Cechosloveniae, 55(1), 44–57.

Dworakowska, I. & Viraktamath, C.A. (1975a) On some Typhlocybinae from India (Auchenorrhyncha, Cicadellidae). Bulletin de l'Academie Polonaise des Sciences, Serie des Sciences Biologiques, 23(8), 521–530.

Dworakowska, I. (1970h) On some genera of Typhlocybini and Empoascini (Auchenorrhyncha, Cicadellidae, Typhlocybinae). Bulletin de l'Academie Polonaise des Sciences, Serie des Sciences Biologiques, 18(11), 707–716.

Dworakowska, I. (1976b) On some Oriental and Ethiopian Typhlocybinae (Homoptera, Auchenorrhyncha, Cicadellidae). Reichenbachia, 16(1), 1–51.

Dworakowska, I. (1977i) On some Typhlocybinae from Vietnam (Homoptera: Cicadellidae). Folia Entomologica Hungarica, 30(2), 9–47.

Husain, M.A. & Lal, K.B. (1940a) The bionomics of Empoasca devastans Distant on some varieties of cotton in the Punjab. Indian Jour. Entomol., 2, 123–136.

Ishida, S. (1913a) Report on the injurious and useful insects of cotton. Govt. of Formosa, Bureau of Industry and Production, Publication No., 12, 1–3.

Jacobi, A.F.V. (1941a) Die Zikadenfauna der Kleinen Sundainseln. Nach der Expeditionsausbeute von B. Rensch. Zoologische Jahrbücher. Abteilung für Systemetik, Ökologie und Geographie der Tiere. Jena, 74, 277–322 + pl. 5.

Khatri, I., Rustamani, M.A., Ahmed, Z., Sultana, R. & Zaidi, S. (2013) Host records of Typhlocybinae (Auchenorrhyncha: Hemiptera: Cicadellidae) of Zoological Museum, University of Karachi, Pakistan. Pakistan Journal of Zoology, 45(5), 1263–1271.

Li, Z.-Z., Xu, P. & Zhang, B. (2005) Homoptera: Hecalinae, Coelidiinae, Iassinae, Typhlocybinae, Nirvaninae, Evacanthinae and Macropsinae. In M.-F. Yang & D.-C. Jin (Eds.). Insect fauna from the National Natural Reserve of Guizhou Province, China. 4. Insects from Dashahe Nature Reserve of Guizhou. Guizhou People Publishing House, Guiyang. pp. 143–152. [in Chinese]

Linnavuori, R.E. (1975c) Homoptera: Cicadellidae, Supplement. Insects of Micronesia, 6(9), 611–632.

Matsumura, S. (1916b) Oyo konchugaku. [Applied Entomology]. Keiseisha Shoten. Tokyo, Vol. 1, 2 + 11 + 731 + 12 + 14 pp. + 50 pls. [in Japanese]

Matsumura, S. (1934b) Eupterygidae. In Catalogue of Japanese insects. Vol. 3, pp. 1–15.

Maxwell-Lefroy, H. (1906a) Indian insect pests. Office Superintendent of Government Printing. Calcutta, 318 pp.

Melichar, L. (1914b) Homopteren von Java, gesammelt von Herrn Edw. Jacobson. Not. Leyden Mus., 36(9), 91–147 + pl. 3.

Merino, G. (1936a) Philippine Cicadellidae (Homoptera). Philippine Journal of Science, 61(3), 307–400, 4 pl.

Metcalf, Z.P. (1968a) General catalogue of the Homoptera. Fascicle VI. Cicadelloidea. Part 17. Cicadellidae. Washington, D.C.: USDA. VII+1513 p.

Schumacher, F. (1915b) "Homoptera." In H. Sauter's Formosa-Ausbeute. Supplementa Entomologica, 4, 108–142.

Shiraki, T. (1913b) Investigations upon insects injurious to cotton. Special Report of the Formosa Agricultural Experiment Station, 5, 1–282. [1912; in Chinese]

Sohi, A.S. (1983a) The oriental Typhlocybinae with special reference to the pests of cotton and rice: a review. Proceedings of the 1st International Workshop on Leafhoppers and Planthoppers of Economic Importance. Commonwealth Institute of Entomology, London, 4-7 Oct. 1982. London. P. 49-74.
Xu, Y., Wang, Y.-R., Dietrich, C.H., Fletcher, M.J. & Qin, D.-Z. (2017a) Review of Chinese species of the leafhopper genus Amrasca Ghauri (Hemiptera, Cicadellidae, Typhlocybinae), with description of a new species, species checklist and notes on the identity of the Indian cotton leafhopper. Zootaxa, 4353(2), 360–370. https://doi.org/10.11646/zootaxa.4353.2.7

Stats

	Valid names
Rank	Extant	Fossil	Invalid	Total
species	1	0	24	25